A few years ago I wrote a post on niche partitioning among herons and egrets. That post was inspired by watching several species of herons and egrets foraging for food along Putah Creek near Davis, CA, and the resource they were partitioning into niches was food.
Recently, as part of my work at the Sacramento-San Joaquin Delta Conservancy, I encountered another example of niche partitioning by herons and egrets. This time, the resource these birds are partitioning into niches is nesting trees.
One of the grants that I manage at the Delta Conservancy is at the Cosumnes River Preserve and it includes and grove of large Valley Oak trees that many herons, egrets, and cormorants use as a rookery (a rookery is a colony of breeding animals, generally birds). One way that the various species have evolved to utilize the same trees, and yet avoid directly competing with each other, is for each species to utilize a different part of each tree to nest in.
Great Blue Herons typically nest on the very tops of the crowns of trees, Great Egrets typically nest only in the upper one-third of the canopy, Snowy Egrets typically nest in the middle one-third of the canopy, and Black-crowned Night-Herons prefer to nest in the lower one-third of the canopy (see the image below).

I think this stratifying of nesting locations is amazing! Species have evolved to fill so many different niches, and so many niches can be divided into finer and finer gradations. I wonder if there is really any limit to how many species can evolve, and how complex an ecosystem can develop, in a give location.
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